School of Natural and Environmental Sciences

Staff Profile

John Knowles

Marine Laboratory Technician/ Boat Crew



M.Phil. in Behavioural Ecology (1999)
B.Sc. (Hons.) Environmental Biology. University of Sunderland (1995).


  • Davies MS, Knowles AJ. Effects of parasitism on the behaviour and ecology of a common marine snail (Littorina littorea(L)). Journal of Experimental Marine Biology and Ecology 2001, 260(2), 155-167.
  • Knowles A. J. & Davies M. S. A novel method to assess the cercariae emission intensity of littorinid populations. Hydrobiologia 378, 237-239 1998.
  • Davies. M. S., Knowles A. J., Edmondston P.& Hutchinson N. The use of a commercial insect trapping compound to maintain grazer densities on rocky shores. Trans. nat. Hist. Soc. Northumb 57, 185-190 1997.